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Metal ion imbalance and oxidative stress trigger the overproduction of Aβ by activating β- or γ-secretases and inhibiting α-secretase, and also cause tau hyperphosphorylation by activating protein kinases, such as glycogen synthase kinase-3β (GSK-3β), cyclin-dependent protein kinase-5 and MAPKs, and inhibiting protein phosphatase 2A (Wang et al., 2020)